Li Sze-Chung
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Li Sizhong (; February 19, 1921 – January 11, 2009) was an
ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish ( Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 33,400 species of fish had been described as of Octobe ...
with the Institute of Zoology () at the
Chinese Academy of Sciences The Chinese Academy of Sciences (CAS); ), known by Academia Sinica in English until the 1980s, is the national academy of the People's Republic of China for natural sciences. It has historical origins in the Academia Sinica during the Republ ...
. Throughout his research career, he made numerous discoveries of new fish
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
(or
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
), and published many books and research papers describing the
fauna Fauna is all of the animal life present in a particular region or time. The corresponding term for plants is ''flora'', and for fungi, it is '' funga''. Flora, fauna, funga and other forms of life are collectively referred to as '' biota''. Zoo ...
and geographical distribution of fishes in China and beyond. He translated and helped publication of the Chinese editions of ''
Fishes of the World ''Fishes of the World'' by the American ichthyologist Joseph S. Nelson (1937–2011) is a standard reference for fish systematics. Now in its fifth edition (2016), the work is a comprehensive overview of the diversity and classification of the ...
'' (2nd edition, by Joseph S. Nelson) and ''Fish Migration'' (a popular science book by Russian zoologist Peter Schmidt). Li was the major author of two published volumes in the Fauna Sinica monograph series, systematically reviewing and describing
orders Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...
of bony fishes that include flat fish,
cod Cod is the common name for the demersal fish genus '' Gadus'', belonging to the family Gadidae. Cod is also used as part of the common name for a number of other fish species, and one species that belongs to genus ''Gadus'' is commonly not call ...
, silverside,
pearlfish Pearlfish are marine fish in the ray-finned fish family Carapidae. Pearlfishes inhabit the tropical waters of the Atlantic, Indian, and Pacific Oceans at depths to , along oceanic shelves and slopes. They are slender, elongated fish with no ...
,
killifish A killifish is any of various oviparous (egg-laying) cyprinodontiform fish (including families Aplocheilidae, Cyprinodontidae, Fundulidae, Profundulidae and Valenciidae). All together, there are 1,270 species of killifish, the biggest family ...
,
flying fish The Exocoetidae are a family of marine fish in the order Beloniformes class Actinopterygii, known colloquially as flying fish or flying cod. About 64 species are grouped in seven to nine genera. While they cannot fly in the same way a bird do ...
, etc. in or near China.Li, S.Z., Zhang, C.G.: ''Fauna Sinica, Osteichthyes: Atheriniformes, Cyprinodontiformes, Beloniformes, Ophidiiformes, Gadiformes'', Science Press, Beijing, 2011. He had written over 40
popular science ''Popular Science'' (also known as ''PopSci'') is an American digital magazine carrying popular science content, which refers to articles for the general reader on science and technology subjects. ''Popular Science'' has won over 58 awards, incl ...
articles about
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
on Chinese newspapers and magazines, and been responsible for compiling and editing fish-related entries in several standard reference books (including
Encyclopedia of China The ''Encyclopedia of China'' () is the first large-entry modern encyclopedia in the Chinese language. The compilation began in 1978. Published by the Encyclopedia of China Publishing House, the encyclopedia was issued one volume at a time, begin ...
and ''Zhonghua Dadian''). His works on freshwater ichthyofauna of China as well as other aspects of ichthyology are considered to be among some of the most notable in the Chinese ichthyology literature. 《20世纪中国学术大典: 生物学》,钱迎倩、王亚辉主编,福建教育出版社,2004
Pages 74 and 237.


Career and research

Right after finishing graduate studies in ichthyology in 1950, Li initially worked on Chinese nomenclature of vertebrates, specializing in fishes, within the publication office of the newly-formed
Chinese Academy of Sciences The Chinese Academy of Sciences (CAS); ), known by Academia Sinica in English until the 1980s, is the national academy of the People's Republic of China for natural sciences. It has historical origins in the Academia Sinica during the Republ ...
. Later in the year, he joined a team of zoologists (including
Tso-hsin Cheng Tso-hsin Cheng (郑作新 also transcribed as Zheng Zuoxin) (18 November 1906 – 27 June 1998) was a Chinese ornithologist known for his seminal work on the birds of China and mentoring a generation of researchers. Educated in the United States, ...
) engaged in consolidating and sorting out zoological specimen inherited from Fan Memorial Institute of Biology and National Academy of Peiping, two predecessors of the Academy of Sciences, in preparation of the later formation of the Institute of Zoology. Within five years
a standard reference on Chinese nomenclature of vertebrates
was published, with a select group of Chinese zoologists as co-authors and contributors. Among them, Li and his mentor Tchang Chun-Lin, along with
Chu Yuan-Ting Chu or CHU may refer to: Chinese history * Chu (state) (c. 1030 BC–223 BC), a state during the Zhou dynasty * Western Chu (206 BC–202 BC), a state founded and ruled by Xiang Yu * Chu Kingdom (Han dynasty) (201 BC–70 AD), a kingdom of the Ha ...
, were responsible for standardization of fish nomenclature in the Chinese language. Starting from the early 1950s, Li helped to initiate and participated in China's systematic surveys of
marine fish Saltwater fish, also called marine fish or sea fish, are fish that live in seawater. Saltwater fish can swim and live alone or in a large group called a school. Saltwater fish are very commonly kept in aquariums for entertainment. Many saltwater f ...
es in
Bohai Sea The Bohai Sea () is a marginal sea approximately in area on the east coast of Mainland China. It is the northwestern and innermost extension of the Yellow Sea, to which it connects to the east via the Bohai Strait. It has a mean depth of a ...
,
Yellow Sea The Yellow Sea is a marginal sea of the Western Pacific Ocean located between mainland China and the Korean Peninsula, and can be considered the northwestern part of the East China Sea. It is one of four seas named after common colour terms ...
and
South China Sea The South China Sea is a marginal sea of the Western Pacific Ocean. It is bounded in the north by the shores of South China (hence the name), in the west by the Indochinese Peninsula, in the east by the islands of Taiwan and northwestern Phil ...
, along China's coast. His scientific career, however, was interrupted in 1957 when he was labelled a "rightist" during the
Anti-Rightist Movement The Anti-Rightist Campaign () in the People's Republic of China, which lasted from 1957 to roughly 1959, was a political campaign to purge alleged "Rightists" within the Chinese Communist Party (CCP) and the country as a whole. The campaign was l ...
. From then until the mid-1970s, while his research and publication capacity had been severely impacted by whims of political campaigns and turmoils, he managed to conduct surveys of
freshwater fish Freshwater fish are those that spend some or all of their lives in fresh water, such as rivers and lakes, with a salinity of less than 1.05%. These environments differ from marine conditions in many ways, especially the difference in levels of s ...
es in the
Yellow River The Yellow River or Huang He (Chinese: , Standard Beijing Mandarin, Mandarin: ''Huáng hé'' ) is the second-longest river in China, after the Yangtze River, and the List of rivers by length, sixth-longest river system in the world at th ...
, Ou River and
Ling River The Jiao River () is a river in Zhejiang Province, China. Its upper stretches are called Yong'an Brook (永安溪) and Ling River (灵江). It flows into the Taizhou Bay, Yueqing Bay and Sanmen Bay of the East China Sea at the Port of Taizhou The ...
, as well as in endorheic regions such as
Xinjiang Xinjiang, SASM/GNC: ''Xinjang''; zh, c=, p=Xīnjiāng; formerly romanized as Sinkiang (, ), officially the Xinjiang Uygur Autonomous Region (XUAR), is an autonomous region of the People's Republic of China (PRC), located in the northwest ...
and
Gansu Gansu (, ; alternately romanized as Kansu) is a province in Northwest China. Its capital and largest city is Lanzhou, in the southeast part of the province. The seventh-largest administrative district by area at , Gansu lies between the Tibet ...
provinces in Western China. While blacklisted as a "rightist" and assigned to work on aquaculture-related research in the early 1960s, he discovered a salmonid fish
serendipitously Serendipity is an unplanned fortunate discovery. Serendipity is a common occurrence throughout the history of product invention and scientific discovery. Etymology The first noted use of "serendipity" was by Horace Walpole on 28 January 1754. I ...
in the vicinity of
Qinling Mountains The Qinling () or Qin Mountains, formerly known as the Nanshan ("Southern Mountains"), are a major east–west mountain range in southern Shaanxi Province, China. The mountains mark the divide between the drainage basins of the Yangtze and Yellow ...
(considered to be a
glacial refugium A glacial refugium (''plural refugia'') is a geographic region which made possible the survival of flora and fauna in times of ice ages and allowed for post-glacial re-colonization. Different types of glacial refugia can be distinguished, namely nun ...
roughly in central part of China), which is later named '' Brachymystax tsinlingensis'' Li, 1966. For a relatively tranquil period of about a year before the
Cultural Revolution The Cultural Revolution, formally known as the Great Proletarian Cultural Revolution, was a sociopolitical movement in the People's Republic of China (PRC) launched by Mao Zedong in 1966, and lasting until his death in 1976. Its stated goal ...
, he was allowed to publish research results related to the salmonid fish, fishes in the Yellow River, and earlier surveys of marine fishes. But only until after
rehabilitation Rehabilitation or Rehab may refer to: Health * Rehabilitation (neuropsychology), therapy to regain or improve neurocognitive function that has been lost or diminished * Rehabilitation (wildlife), treatment of injured wildlife so they can be retur ...
in 1978, Li was able to resume normal research activities. In his book ''Studies on zoogeographical divisions for fresh water fishes of China'', Li divided the fauna of freshwater fishes in China into five major regions based on characteristics of fish
species distribution Species distribution —or species dispersion — is the manner in which a biological taxon is spatially arranged. The geographic limits of a particular taxon's distribution is its range, often represented as shaded areas on a map. Patterns of ...
s, geographic environments and geological histories of these regions. The five freshwater regions in China are: (1) Northern region containing upper reaches of rivers that flow into the
Arctic Ocean The Arctic Ocean is the smallest and shallowest of the world's five major oceans. It spans an area of approximately and is known as the coldest of all the oceans. The International Hydrographic Organization (IHO) recognizes it as an ocean, a ...
, as well as
Amur River The Amur (russian: река́ Аму́р, ), or Heilong Jiang (, "Black Dragon River", ), is the world's List of longest rivers, tenth longest river, forming the border between the Russian Far East and Northeast China, Northeastern China (Inne ...
and its tributaries; (2) Western China region; (3)
Mongolian Plateau The Mongolian Plateau is the part of the Central Asian Plateau lying between 37°46′-53°08′N and 87°40′-122°15′E and having an area of approximately . It is bounded by the Greater Hinggan Mountains in the east, the Yin Mountains to th ...
; (4) Eastern China region; and (5) Southern China region. According to his method, the demarcation line between
Holarctic The Holarctic realm is a biogeographic realm that comprises the majority of habitats found throughout the continents in the Northern Hemisphere. It corresponds to the floristic Boreal Kingdom. It includes both the Nearctic zoogeographical reg ...
and
Indomalayan realm The Indomalayan realm is one of the eight biogeographic realms. It extends across most of South and Southeast Asia and into the southern parts of East Asia. Also called the Oriental realm by biogeographers, Indomalaya spreads all over the Indi ...
s in China, as far as freshwater fishes are concerned, lies largely along the
Himalayas The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 100 ...
and
Nanling Mountains The Nanling (), also known as the Wuling (), is a major mountain range in Southern China that separates the Pearl River Basin from the Yangtze Valley and serves as the dividing line between south and central subtropical zones. The main range ...
ranges, where the Southern China region is basically on the south side of the Nanling Mountains, a
drainage divide A drainage divide, water divide, ridgeline, watershed, water parting or height of land is elevated terrain that separates neighboring drainage basins. On rugged land, the divide lies along topographical ridges, and may be in the form of a singl ...
between the
Yangtze River The Yangtze or Yangzi ( or ; ) is the longest list of rivers of Asia, river in Asia, the list of rivers by length, third-longest in the world, and the longest in the world to flow entirely within one country. It rises at Jari Hill in th ...
and the
Pearl River The Pearl River, also known by its Chinese name Zhujiang or Zhu Jiang in Mandarin pinyin or Chu Kiang and formerly often known as the , is an extensive river system in southern China. The name "Pearl River" is also often used as a catch-a ...
. This idea is close to those of
Wallace Wallace may refer to: People * Clan Wallace in Scotland * Wallace (given name) * Wallace (surname) * Wallace (footballer, born 1986), full name Wallace Fernando Pereira, Brazilian football left-back * Wallace (footballer, born 1987), full name ...
and
Mori Mori is a Japanese and Italian surname, and also a Persian pet name for Morteza. It is also the name of two clans in Japan, and one clan in India. Italian surname *Barbara Mori, Uruguayan-Mexican actress * Camilo Mori, Chilean painter * Cesare ...
, and contrasts with the prevalent view of demarcation line for
terrestrial animals Terrestrial animals are animals that live predominantly or entirely on land (e.g. cats, dogs, ants, spiders), as compared with aquatic animals, which live predominantly or entirely in the water (e.g. fish, lobsters, octopuses), and amphibians, ...
between Holarctic and Indomalaya regions along the
Qinling Mountains The Qinling () or Qin Mountains, formerly known as the Nanshan ("Southern Mountains"), are a major east–west mountain range in southern Shaanxi Province, China. The mountains mark the divide between the drainage basins of the Yangtze and Yellow ...
(known as
Qinling–Huaihe Line The Qinling–Huaihe Line () is a reference line used by geographers to distinguish between northern and southern China, corresponding roughly to the 33rd parallel. Qinling refers to the Qin Mountains, and Huaihe refers to the Huai River. Runn ...
, though the Qinling Mountains does not extend into Eastern part of China). Freshwater Ecoregions of the World (FEOW), a collaborative global
biodiversity Biodiversity or biological diversity is the variety and variability of life on Earth. Biodiversity is a measure of variation at the genetic (''genetic variability''), species (''species diversity''), and ecosystem (''ecosystem diversity'') l ...
project partly sponsored by
World Wide Fund for Nature The World Wide Fund for Nature Inc. (WWF) is an international non-governmental organization founded in 1961 that works in the field of wilderness preservation and the reduction of human impact on the environment. It was formerly named the Wor ...
(WWF), cites this book as a source of references in delineation of
freshwater ecoregion An ecoregion (ecological region) or ecozone (ecological zone) is an ecologically and geographically defined area that is smaller than a bioregion, which in turn is smaller than a biogeographic realm. Ecoregions cover relatively large areas of l ...
s of China. Two of his books are published posthumously. In ''Fishes of the Yellow River and Beyond'', he described over 170 native fish species and the characteristics of fish distribution in the Yellow River and its tributaries; a collection of his papers and popular science articles, as well as a personal memoir, are also included in the posthumous book published in Taiwan; this book was re-published in the mainland in 2017. Comparing fish species in the Yellow River in 2010–2015 and the survey results originally reported by Li in 1965 as part of his research on the book, two recent Chinese studies conclude that only about half of the native fishes in the Yellow River could still be found, due to anthropogenic environmental alterations and increased presence of
introduced species An introduced species, alien species, exotic species, adventive species, immigrant species, foreign species, non-indigenous species, or non-native species is a species living outside its native distributional range, but which has arrived there ...
over the past half-century. Li was also the primary contributor to a volume of the ''Fauna Sinica'' series covering the
orders Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...
of
Atheriniformes The Atheriniformes, also known as the silversides, are an order of ray-finned fishes that includes the Old World silversides and several less-familiar families, including the unusual Phallostethidae. The order includes at least 354 species. T ...
,
Cyprinodontiformes Cyprinodontiformes is an order of ray-finned fish, comprising mostly small, freshwater fish. Many popular aquarium fish, such as killifish and live-bearers, are included. They are closely related to the Atheriniformes and are occasionally inc ...
,
Beloniformes Beloniformes is an order composed of six families (and about 264 species) of freshwater and marine ray-finned fish: * Adrianichthyidae (ricefish and medakas) * Belonidae (needlefish) * Exocoetidae (flyingfishes) * Hemiramphidae (halfbeaks) ...
,
Ophidiiformes Ophidiiformes is an order of ray-finned fish that includes the cusk-eels (family Ophidiidae), pearlfishes (family Carapidae), viviparous brotulas (family Bythitidae), and others. Members of this order have small heads and long slender bodies. Th ...
and
Gadiformes Gadiformes are an order of ray-finned fish, also called the Anacanthini, that includes the cod. Many major food fish are in this order. They are found in marine waters throughout the world and the vast majority of the species are found in tempe ...
, which was published posthumously in 2011. Another volume of the ''Fauna Sinica'' series which he worked on during his last years, covering fish species in China within the orders of
Beryciformes The Beryciformes are a poorly-understood order of carnivorous ray-finned fishes consisting of 7 families, 30 genera, and 161 species. They feed on small fish and invertebrates. Beyond this, little is known about the biology of most member specie ...
,
Zeiformes The Zeiformes are a small order of marine ray-finned fishes most notable for the dories, a group of common food fish. The order consists of about 33 species in seven families, mostly deep-sea types. Zeiform bodies are usually thin and deep. Mo ...
,
Lampriformes Lampriformes is an order of ray-finned fish. Members are collectively called lamprids (which is more properly used for the Lampridae) or lampriforms, and unite such open-ocean and partially deep-sea Teleostei as the crestfishes, oarfish, opah ...
,
Gasterosteiformes Gasterosteoidei is a suborder of ray-finned fishes that includes the sticklebacks and relatives, the 5th edition of '' Fishes of the World'' classifies this suborder within the order Scorpaeniformes. Systematics Gasterosteoidei is treated as ...
,
Mugiliformes The mullets or grey mullets are a family (Mugilidae) of ray-finned fish found worldwide in coastal temperate and tropical waters, and some species in fresh water. Mullets have served as an important source of food in Mediterranean Europe since R ...
and Synbranchiformes (such as
soldierfish Myripristinae is a subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus Genus ( plural genera ) is a taxonomic rank us ...
,
dory A dory is a small, shallow-draft boat, about long. It is usually a lightweight boat with high sides, a flat bottom and sharp bows. It is easy to build because of its simple lines. For centuries, the dory has been used as a traditional fishin ...
,
opah Opahs, also commonly known as moonfish, sunfish (not to be confused with Molidae), kingfish, redfin ocean pan are large, colorful, deep-bodied pelagic lampriform fishes comprising the small family Lampridae (also spelled Lamprididae). The famil ...
,
ribbonfish The ribbonfish are any lampriform fishes in the family Trachipteridae. These pelagic fish are named for their slim, ribbon-like appearance. They are rarely seen alive, as they typically live in deep waters, though are not bottom feeders. The p ...
, stickleback, mullet,
swamp eel The swamp eels (also written "swamp-eels") are a family (Synbranchidae) of freshwater eel-like fishes of the tropics and subtropics. Most species are able to breathe air and typically live in marshes, ponds and damp places, sometimes burying the ...
, etc.), is yet to be published.中国动物志出版信息: Osteichthyes: Beryciformes-Synbranchiformes
/ref>


Fish species discovered or named


Footnotes

{{DEFAULTSORT:Li, Sizhong 1921 births 2009 deaths Chinese ichthyologists Biogeographers Writers from Xinxiang People's Republic of China science writers Victims of the Cultural Revolution Biologists from Henan Beijing Normal University alumni People of the Republic of China People from Huixian 20th-century Chinese zoologists English–Chinese_translators Russian–Chinese_translators